HANOVER — Jaden Craig threw for 311 yards and a touchdown, Charles DePrima rushed for two fourth-quarter touchdowns and Harvard beat Dartmouth, 31-27, on Saturday for its fifth straight win, knocking the Big Green from the ranks of the unbeaten.

It was the lone FCS contest on Saturday featuring two teams with one loss or fewer on the season. The winning streak for Harvard (6-1, 3-1 Ivy League) is its longest since the 2015 squad opened the campaign with eight consecutive victories.

DePrima pulled Harvard within 27-24 with 5:59 remaining in the fourth quarter after capping a 14-play, 94-yard drive with a 6-yard TD. After Dartmouth went three-and-out, Harvard chewed up 3 minutes, 32 seconds of clock tie, ending in a 9-yard touchdown run by DePrima with just 27 seconds left.

The Crimson won at Dartmouth (6-1, 3-1) for the 14th time in 15 meetings since the start of 1994, including a 28-13 decision in 2022.

DePrima opened the scoring on Harvard’s first possession of the game, and Kieran Corr added a 32-yard field goal before Dartmouth scored the next 20 points for a 10-point lead late in the third quarter.

The end of the third quarter saw 21 points scored in the final 3:03. Dartmouth quarterback Jackson Proctor connected with Chris Corbo from 11 yards out and Daniel Haughton on a 72-yard catch-and-run sandwiched around Craig’s 29-yard TD pass to Seamus Gilmartin.

Cooper Barkate led Harvard’s receivers with 124 yards.

Proctor passed for 235 yards and two touchdowns for Dartmouth.

Penn 38, Brown 28

PROVIDENCE, R.I. — Liam O’Brien threw for two touchdowns and ran for another to lead Pennsylvania to its first Ivy League win this season.

O’Brien was 18-of-24 passing for 247 yards with touchdown throws to Jared Richardson and Justin Cayenne. His 10-yard TD run in the final minute of the first half gave the Quakers (3-4, 1-3) a 21-7 lead.

Malachi Hosley added to the lead early in the third quarter with an 11-yard TD run.

Cornell 49, Princeton 35

PRINCETON, N.J. — Jameson Wang threw a pair of touchdown passes and ran for a third in a 21-0 first quarter to kick-start Cornell to a win over of Princeton in an Ivy League battle on Saturday.

The Big Red (3-4, 2-2), who won for just the 38th time in the 106-game history of the series, raced to a 28-0 lead before the Tigers (2-5, 1-3) could find the end zone.

FRIDAY’S GAMEColumbia 13, Yale 10

NEW YORK — Cole Freeman threw a go-ahead 44-yard touchdown pass to Bryson Canty with 4:39 left to play and Columbia pulled out a victory over Yale.